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Tribe Cacteae: Ferocactus Clade
Family: Cactaceae
Subfamily: Cactoideae
Tribe: Cacteae
Clade: Ferocactus
The Ferocactus clade contains to following Species (alphabetical order) according to Butterworth's paper (see bottom of page).
Echinocactus grusonii
Ferocactus cylindraceus
Ferocactus flavovirens
Ferocactus glaucescens
Ferocactus latispinus
Ferocactus robustus
Ferocactus wislizenii
Glandulicactus crassihamatus
Glandulicactus uncinatus
Leuchtenbergia principis
Thelocactus conothelos
Thelocactus hastifer
Thelocactus macdowellii
This page is put together from the paper by Charles A. Butterworth, J. Hugo Cota-Sanchez, and Robert S. Wallace.
Molecular Systematics of Tribe Cacteae (Cactaceae: Cactoideae): A Phylogeny Based on rpl16 Intron Sequence Variation
Abstract Parsimony analysis of plastid rpl16 sequences from 62 members of Tribe Cacteae, and four outgroup taxa yielded 1296 equally parsimonious trees of length 666. Strict consensus evaluation of these trees established a highly pectinate topology, which delimited clades within the tribe that correspond to several previously considered generic groups. Aztekium and Geohintonia, which manifest ribs in their stem morphology were shown to represent an early divergence in the tribe, forming a sister group to remaining members of the tribe. Clades containing other genera having ribbed stems also are basal to those that develop tubercles. The most derived clade forms a distinct group of typically small stemmed species with tubercular stem morphology. Within Mammillaria, species formerly placed in the genus Cochemiea and members of the Series Ancistracanthae formed a well-supported, sister clade to the remaining members of Mammillaria. Length variation of the intron in two members of Mammillaria series Stylothelae was also observed.
